Yale University School of Forestry & Environmental Studies
FES519B - Methods of Ecosystem Analysis 1998
SASSAFRAS
ALLOMETRY
at
TOTOKET MOUNTAIN,
NORTH BRANFORD, CT
1998
 
                ·        

About this Web page
        The data presented below are the result of a term project for Professor Tom Siccama's FES 519b: Methods of Ecosystem Analysis class at the Yale University School of Forestry and Environmental Studies.  Field sampling was done in February-March 1998 at Totoket Mountain near North Branford, CT.  We are indebted to the Regional Water Authority for providing access to the site.
         The allometry equations presented are intended to be used to calculate dry weight, above ground biomass of sassafras trees in winter, without leaves.

 
What's below
        On this page, you'll find the following information about the Sassafras allometry project at Totoket Mountain:


What is allometry?
 
        Allometry is all about studying the relative sizes of plant parts.  Usually, relationships
between dbh (diameter at breast height, or 1.37 m up from ground level), tree height, total biomass, leaf weight, etc., are calculated.  For example, what we do here is prepare equations (regressions) to calculate the total above-ground biomass of a Sassafras tree in winter (without leaves) as a function of dbh: we are therefore implicitly assuming that biomass is directly related to tree diameter.

        Allometry is useful because it allows the total biomass of a forest or stand to be estimated, without having to cut down all the trees, take them back to the lab, dry the pieces in an oven, and then weigh all the pieces.   Part of our study at Totoket Mountain was to estimate the biomass of the entire forest: click here to jump to that page.


How do we do it, anyway?
 

         To calculate the total biomass of our Sassafras trees, we had to calculate

          In case you want to try this at home, here's what we did to each tree!  We did it for four trees, of different sizes and shapes, so that our results would be more applicable to "all" sassafras trees.

        By combining our estimates of bole and branch weights, we calculated the total above-ground biomass of each tree.  We then estimated another regression, this one relating tree diameter to total above ground biomass.

        When we calculated our regression equations, we used a log-log transformation on the data.  Rather than make total above ground biomass a function of dbh, we made log10 (total above ground biomass) a function of log10 (dbh).  The relationship between the transformed variables is a linear one: we can calculate this regression line much more easily than if we dealt with untransformed data.  Another effect that the log transformation has is that it can make large absolute differences appear relatively small: for example, although the difference between 600 kg and 1000 kg is 400 kg, log(600) = 2.78, log (1000) = 3, and the difference of the logs, 0.22, is less than 8%!

        We arrived at the following equations:

        Bole biomass:
         log10 (bole biomass, g) = 2.3904 * log10 (dbh, cm) + 1.8632

        Individual branch weight:
        log10 (branch dry weight, g) = 2.807 * log10 (branch diameter, cm) + 1.4418

        Total branch weight:
        log10 (total branch biomass, g) = 2.5533 * log10 (dbh, cm) + .9020

        Total above-ground biomass
         log10 (bole + branch biomass, g) = 2.3836 * log10 (dbh, cm) + 1.9566
 



Basic Tree Data

As described above, we cut down four sassafras trees on the east slope at Totoket Mountain, North Branford, CT for analysis.  Basic data for these four trees are given below.
 
Tree 1
Tree 2
Tree 3
Tree 4
Total height, m
9.1
16.4
14.7
17.2
Tree age (at base), years
73.0
77.0
77.0
Avg. annual radial increment, mm, last 5 years
2.0
0.6
0.9
DBH, cm
5.0
23.4
11.8
16.7
Diameter at base, cm
6.4
28.1
13.9
21.1
Crown width, widest, m
2.0
6.3
3.7
2.6
Crown width, narrowest, m
1.5
5.9
3.5
2.0
Height to lowest live branch, m
6.8
7.2
8.0
7.0
Estimated bole dry weight, OD kg
4.7
137.3
27.5
71.2
Estimated total branch dry weight, OD kg 
       (live + dead)
0.6
32.0
3.2
5.6
Weight dead branches, OD kg
0.3
4.9
1.2
2.2
Estimated biomass, OD kg 
       (above ground, no leaves)
5.4
169.3
30.7
76.7
Average bark thickness, cm
2.1
1.1
1.5
Bark water content (g H20 / g OD)
31%
26%
30%
Wood water content (g H20 / g OD)
59%
39%
45%
Bark density, g / cc
0.26
0.30
0.24
Wood density, g/cc
0.42
0.41
0.41
Total bark weight in bole, OD kg
19.4
4.7
10.1
Total wood weight in bole, OD kg
117.0
22.8
61.1
Note: some data not available for tree 1, because no cookies were taken (the entire tree was brought back to the lab, where the branches and bole were separated and oven-dried).
 

        We estimated height-age curves for Trees 2, 3, and 4 by aging the cookies taken at 1 m intervals.  In some cases, rot inside the bole prevented an accurate measurement, because rings could not be counted all the way back to age 1.  Best-fit curves (third order polynomial equations) suggest that, in the case of Trees 2 and 3, some sort of release has occurred during the last 20 or so years.

           Tree 2

            Tree 3

             Tree 4
 

      We also used data collected by previous years' FES 519b classes.  This and more can be found in our downloadable data workbook.


Estimating total bole weight
 
Click here to see the data for this graph.
 


Estimating bole volume

        The bole of a tree can be considered as any of a number of three-dimensional solids.  Usually, the bole volume is estimated as if it were a cone, a paraboloid, or a neolid (a solid that flares at the base).  We found that sassafras is pretty much cone-shaped, as the following graph illustrates.


Estimating weight of individual live branches
  
Click here to see the data for this graph.
 


Estimating weight of all branches, live and dead
 
Click here to see the data for this graph.


Estimating above-ground biomass of whole tree

Click here to see the data for this graph.

The graph below is the same as the graph above, except that the log-log transformation has not been carried out.  Observe that the best-fit line is no longer linear.  Also, note that even though it fits the data very well, the biomass predicted by the equation is rarely exactly what we measured in the lab (the deviation between measured and calculated biomass is greater than 20% for some of the larger diameter trees)!
 
 



How fast are these trees growing?

        We measured the average radial growth over the last five years (1996-1992: the 1997 ring was usually impossible to read due to the bark having been removed) for each tree from each cookie (1 m height increments along the bole).  Tree 2 appears to have put on more radial growth high in the bole than low in the bole; for Trees 3 and 4, the radial growth changes little with height.

 
By taking the average annual radial growth,  multiplying this by five, doubling that (to get the diameter change over the last five years), and then subtracting this value from the measured dbh, we estimated the tree's diameter in 1993.  We then put this value into our biomass equation, and approximated the tree's biomass in 1993.  The trees appear to have grown by 25-30% over the last five years, as the following table shows.
 
1998 Biomass 
(from lab, kg)
1993 dbh 
(cm)
1998 dbh 
(cm)
1993 Biomass 
(from eq., kg)
1998 Biomass
(from eq., kg)
% Change
Tree 2
169.3
21.4
23.4
110.5
136.8
+24%
Tree 3
30.7
10.6
11.8
20.6
26.6
+29%
Tree 4
76.7
14.9
16.7
46.5
61.1
+31%
 


Tell me more about Sassafras
        Sassafras albidum is a ....
        Here are some links to other Web pages about Sassafras:

Download our data sheets

        If you want to use our data and see our calculations, you can download the Microsoft Excel workbook.  Click here to download.
 


 
Sassafras allometry brought to you by 
    Jocelyn Forbush 
    Andrew Richardson 
    Yale School of Forestry & Environmental Studies 

With thanks to 
    Regional Water Authority 
    Previous FES 519b sassafras allometry groups 
    Tom Siccama 
    Dan Vogt 
    All the other Crooked Brookers